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¡¡be¡¡seen£»¡¡the¡¡linking¡¡of¡¡diverse¡¡families¡¡through¡¡intermediate¡¡ones£»¡¡and¡¡in¡¡particular¡¡with¡¡the¡¡predominance¡¡of¡¡low¡¡types¡¡of¡¡life¡¡in¡¡the¡¡earlier¡¡geological¡¡strata¡£¡¡¡¡Called¡¡upon¡¡constantly¡¡to¡¡classify¡¡the¡¡various¡¡forms¡¡of¡¡life¡¡in¡¡the¡¡course¡¡of¡¡his¡¡systematic¡¡writings£»¡¡he¡¡found¡¡it¡¡more¡¡and¡¡more¡¡difficult¡¡to¡¡draw¡¡sharp¡¡lines¡¡of¡¡demarcation£»¡¡and¡¡at¡¡last¡¡the¡¡suspicion¡¡long¡¡harbored¡¡grew¡¡into¡¡a¡¡settled¡¡conviction¡¡that¡¡there¡¡is¡¡really¡¡no¡¡such¡¡thing¡¡as¡¡a¡¡species¡¡of¡¡organism¡¡in¡¡nature£»¡¡that¡¡¡¨species¡¨¡¡is¡¡a¡¡figment¡¡of¡¡the¡¡human¡¡imagination£»¡¡whereas¡¡in¡¡nature¡¡there¡¡are¡¡only¡¡individuals¡£

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In¡¡Philosophie¡¡Zoologique¡¡£¨1809£©£»¡¡Lamarck¡¡first¡¡explicitly¡¡formulated¡¡his¡¡ideas¡¡as¡¡to¡¡the¡¡transmutation¡¡of¡¡species£»¡¡though¡¡he¡¡had¡¡outlined¡¡them¡¡as¡¡early¡¡as¡¡1801¡£¡¡¡¡In¡¡this¡¡memorable¡¡publication¡¡not¡¡only¡¡did¡¡he¡¡state¡¡his¡¡belief¡¡more¡¡explicitly¡¡and¡¡in¡¡fuller¡¡detail¡¡than¡¡the¡¡idea¡¡had¡¡been¡¡expressed¡¡by¡¡any¡¡predecessor£»¡¡but¡¡he¡¡took¡¡another¡¡long¡¡forward¡¡step£»¡¡carrying¡¡him¡¡far¡¡beyond¡¡all¡¡his¡¡forerunners¡¡except¡¡Darwin£»¡¡in¡¡that¡¡he¡¡made¡¡an¡¡attempt¡¡to¡¡explain¡¡the¡¡way¡¡in¡¡which¡¡the¡¡transmutation¡¡of¡¡species¡¡had¡¡been¡¡brought¡¡about¡£¡¡The¡¡changes¡¡have¡¡been¡¡wrought£»¡¡he¡¡said£»¡¡through¡¡the¡¡unceasing¡¡efforts¡¡of¡¡each¡¡organism¡¡to¡¡meet¡¡the¡¡needs¡¡imposed¡¡upon¡¡it¡¡by¡¡its¡¡environment¡£¡¡Constant¡¡striving¡¡means¡¡the¡¡constant¡¡use¡¡of¡¡certain¡¡organs¡£¡¡Thus¡¡a¡¡bird¡¡running¡¡by¡¡the¡¡seashore¡¡is¡¡constantly¡¡tempted¡¡to¡¡wade¡¡deeper¡¡and¡¡deeper¡¡in¡¡pursuit¡¡of¡¡food£»¡¡its¡¡incessant¡¡efforts¡¡tend¡¡to¡¡develop¡¡its¡¡legs£»¡¡in¡¡accordance¡¡with¡¡the¡¡observed¡¡principle¡¡that¡¡the¡¡use¡¡of¡¡any¡¡organ¡¡tends¡¡to¡¡strengthen¡¡and¡¡develop¡¡it¡£¡¡But¡¡such¡¡slightly¡¡increased¡¡development¡¡of¡¡the¡¡legs¡¡is¡¡transmitted¡¡to¡¡the¡¡off¡¡spring¡¡of¡¡the¡¡bird£»¡¡which¡¡in¡¡turn¡¡develops¡¡its¡¡already¡¡improved¡¡legs¡¡by¡¡its¡¡individual¡¡efforts£»¡¡and¡¡transmits¡¡the¡¡improved¡¡tendency¡£¡¡Generation¡¡after¡¡generation¡¡this¡¡is¡¡repeated£»¡¡until¡¡the¡¡sum¡¡of¡¡the¡¡infinitesimal¡¡variations£»¡¡all¡¡in¡¡the¡¡same¡¡direction£»¡¡results¡¡in¡¡the¡¡production¡¡of¡¡the¡¡long¡­legged¡¡wading¡­bird¡£¡¡In¡¡a¡¡similar¡¡way£»¡¡through¡¡individual¡¡effort¡¡and¡¡transmitted¡¡tendency£»¡¡all¡¡the¡¡diversified¡¡organs¡¡of¡¡all¡¡creatures¡¡have¡¡been¡¡developedthe¡¡fin¡¡of¡¡the¡¡fish£»¡¡the¡¡wing¡¡of¡¡the¡¡bird£»¡¡the¡¡hand¡¡of¡¡man£»¡¡nay£»¡¡more£»¡¡the¡¡fish¡¡itself£»¡¡the¡¡bird£»¡¡the¡¡man£»¡¡even¡£¡¡¡¡Collectively¡¡the¡¡organs¡¡make¡¡up¡¡the¡¡entire¡¡organism£»¡¡and¡¡what¡¡is¡¡true¡¡of¡¡the¡¡individual¡¡organs¡¡must¡¡be¡¡true¡¡also¡¡of¡¡their¡¡ensemble£»¡¡the¡¡living¡¡being¡£

Whatever¡¡might¡¡be¡¡thought¡¡of¡¡Lamarck's¡¡explanation¡¡of¡¡the¡¡cause¡¡of¡¡transmutationwhich¡¡really¡¡was¡¡that¡¡already¡¡suggested¡¡by¡¡Erasmus¡¡Darwinthe¡¡idea¡¡of¡¡the¡¡evolution¡¡for¡¡which¡¡he¡¡contended¡¡was¡¡but¡¡the¡¡logical¡¡extension¡¡of¡¡the¡¡conception¡¡that¡¡American¡¡animals¡¡are¡¡the¡¡modified¡¡and¡¡degenerated¡¡descendants¡¡of¡¡European¡¡animals¡£¡¡But¡¡people¡¡as¡¡a¡¡rule¡¡are¡¡little¡¡prone¡¡to¡¡follow¡¡ideas¡¡to¡¡their¡¡logical¡¡conclusions£»¡¡and¡¡in¡¡this¡¡case¡¡the¡¡conclusions¡¡were¡¡so¡¡utterly¡¡opposed¡¡to¡¡the¡¡proximal¡¡bearings¡¡of¡¡the¡¡idea¡¡that¡¡the¡¡whole¡¡thinking¡¡world¡¡repudiated¡¡them¡¡with¡¡acclaim¡£¡¡The¡¡very¡¡persons¡¡who¡¡had¡¡most¡¡eagerly¡¡accepted¡¡the¡¡idea¡¡of¡¡transmutation¡¡of¡¡European¡¡species¡¡into¡¡American¡¡species£»¡¡and¡¡similar¡¡limited¡¡variations¡¡through¡¡changed¡¡environment£»¡¡because¡¡of¡¡the¡¡relief¡¡thus¡¡given¡¡the¡¡otherwise¡¡overcrowded¡¡ark£»¡¡were¡¡now¡¡foremost¡¡in¡¡denouncing¡¡such¡¡an¡¡extension¡¡of¡¡the¡¡doctrine¡¡of¡¡transmutation¡¡as¡¡Lamarck¡¡proposed¡£

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Cuvier's¡¡reasoning¡¡has¡¡such¡¡great¡¡historical¡¡interestbeing¡¡the¡¡argument¡¡of¡¡the¡¡greatest¡¡opponent¡¡of¡¡evolution¡¡of¡¡that¡¡daythat¡¡we¡¡quote¡¡it¡¡at¡¡some¡¡length¡£

¡¨The¡¡following¡¡objections£»¡¨¡¡he¡¡says£»¡¡¡¨have¡¡already¡¡been¡¡started¡¡against¡¡my¡¡conclusions¡£¡¡¡¡Why¡¡may¡¡not¡¡the¡¡presently¡¡existing¡¡races¡¡of¡¡mammiferous¡¡land¡¡quadrupeds¡¡be¡¡mere¡¡modifications¡¡or¡¡varieties¡¡of¡¡those¡¡ancient¡¡races¡¡which¡¡we¡¡now¡¡find¡¡in¡¡the¡¡fossil¡¡state£»¡¡which¡¡modifications¡¡may¡¡have¡¡been¡¡produced¡¡by¡¡change¡¡of¡¡climate¡¡and¡¡other¡¡local¡¡circumstances£»¡¡and¡¡since¡¡raised¡¡to¡¡the¡¡present¡¡excessive¡¡difference¡¡by¡¡the¡¡operations¡¡of¡¡similar¡¡causes¡¡during¡¡a¡¡long¡¡period¡¡of¡¡ages£¿

¡¨This¡¡objection¡¡may¡¡appear¡¡strong¡¡to¡¡those¡¡who¡¡believe¡¡in¡¡the¡¡indefinite¡¡possibility¡¡of¡¡change¡¡of¡¡form¡¡in¡¡organized¡¡bodies£»¡¡and¡¡think¡¡that£»¡¡during¡¡a¡¡succession¡¡of¡¡ages¡¡and¡¡by¡¡alterations¡¡of¡¡habitudes£»¡¡all¡¡the¡¡species¡¡may¡¡change¡¡into¡¡one¡¡another£»¡¡or¡¡one¡¡of¡¡them¡¡give¡¡birth¡¡to¡¡all¡¡the¡¡rest¡£¡¡Yet¡¡to¡¡these¡¡persons¡¡the¡¡following¡¡answer¡¡may¡¡be¡¡given¡¡from¡¡their¡¡own¡¡system£º¡¡If¡¡the¡¡species¡¡have¡¡changed¡¡by¡¡degrees£»¡¡as¡¡they¡¡assume£»¡¡we¡¡ought¡¡to¡¡find¡¡traces¡¡of¡¡this¡¡gradual¡¡modification¡£¡¡¡¡Thus£»¡¡between¡¡the¡¡palaeotherium¡¡and¡¡the¡¡species¡¡of¡¡our¡¡own¡¡day£»¡¡we¡¡should¡¡be¡¡able¡¡to¡¡discover¡¡some¡¡intermediate¡¡forms£»¡¡and¡¡yet¡¡no¡¡such¡¡discovery¡¡has¡¡ever¡¡been¡¡made¡£¡¡Since¡¡the¡¡bowels¡¡of¡¡the¡¡earth¡¡have¡¡not¡¡preserved¡¡monuments¡¡of¡¡this¡¡strange¡¡genealogy£»¡¡we¡¡have¡¡no¡¡right¡¡to¡¡conclude¡¡that¡¡the¡¡ancient¡¡and¡¡now¡¡extinct¡¡species¡¡were¡¡as¡¡permanent¡¡in¡¡their¡¡forms¡¡and¡¡characters¡¡as¡¡those¡¡which¡¡exist¡¡at¡¡present£»¡¡or£»¡¡at¡¡least£»¡¡that¡¡the¡¡catastrophe¡¡which¡¡destroyed¡¡them¡¡did¡¡not¡¡leave¡¡sufficient¡¡time¡¡for¡¡the¡¡productions¡¡of¡¡the¡¡changes¡¡that¡¡are¡¡alleged¡¡to¡¡have¡¡taken¡¡place¡£

¡¨In¡¡order¡¡to¡¡reply¡¡to¡¡those¡¡naturalists¡¡who¡¡acknowledge¡¡that¡¡the¡¡varieties¡¡of¡¡animals¡¡are¡¡restrained¡¡by¡¡nature¡¡within¡¡certain¡¡limits£»¡¡it¡¡would¡¡be¡¡necessary¡¡to¡¡examine¡¡how¡¡far¡¡these¡¡limits¡¡extend¡£¡¡This¡¡is¡¡a¡¡very¡¡curious¡¡inquiry£»¡¡and¡¡in¡¡itself¡¡exceedingly¡¡interesting¡¡under¡¡a¡¡variety¡¡of¡¡relations£»¡¡but¡¡has¡¡been¡¡hitherto¡¡very¡¡little¡¡attended¡¡to¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£

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¡¨Nature¡¡appears¡¡also¡¡to¡¡have¡¡guarded¡¡against¡¡the¡¡alterations¡¡of¡¡species¡¡which¡¡might¡¡proceed¡¡from¡¡mixture¡¡of¡¡breeds¡¡by¡¡influencing¡¡the¡¡various¡¡species¡¡of¡¡animals¡¡with¡¡mutual¡¡aversion¡¡from¡¡one¡¡another¡£¡¡Hence¡¡all¡¡the¡¡cunning¡¡and¡¡all¡¡the¡¡force¡¡that¡¡man¡¡is¡¡able¡¡to¡¡exert¡¡is¡¡necessary¡¡to¡¡accomplish¡¡such¡¡unions£»¡¡even¡¡between¡¡species¡¡that¡¡have¡¡the¡¡nearest¡¡resemblances¡£¡¡¡¡And¡¡when¡¡the¡¡mule¡¡breeds¡¡that¡¡are¡¡thus¡¡produced¡¡by¡¡these¡¡forced¡¡conjunctions¡¡happen¡¡to¡¡be¡¡fruitful£»¡¡which¡¡is¡¡seldom¡¡the¡¡case£»¡¡this¡¡fecundity¡¡never¡¡continues¡¡beyond¡¡a¡¡few¡¡generations£»¡¡and¡¡would¡¡not¡¡probably¡¡proceed¡¡so¡¡far¡¡without¡¡a¡¡continuance¡¡of¡¡the¡¡same¡¡cares¡¡which¡¡excited¡¡it¡¡at¡¡first¡£¡¡Thus¡¡we¡¡never¡¡see¡¡in¡¡a¡¡wild¡¡state¡¡intermediate¡¡productions¡¡between¡¡the¡¡hare¡¡and¡¡the¡¡rabbit£»¡¡between¡¡the¡¡stag¡¡and¡¡the¡¡doe£»¡¡or¡¡between¡¡the¡¡marten¡¡and¡¡the¡¡weasel¡£¡¡¡¡But¡¡the¡¡power¡¡of¡¡man¡¡changes¡¡this¡¡established¡¡order£»¡¡and¡¡continues¡¡to¡¡produce¡¡all¡¡these¡¡intermixtures¡¡of¡¡which¡¡the¡¡various¡¡species¡¡are¡¡susceptible£»¡¡but¡¡which¡¡they¡¡would¡¡never¡¡produce¡¡if¡¡left¡¡to¡¡themselves¡£

¡¨The¡¡degrees¡¡of¡¡these¡¡variations¡¡are¡¡proportional¡¡to¡¡the¡¡intensity¡¡of¡¡the¡¡causes¡¡that¡¡produced¡¡themnamely£»¡¡the¡¡slavery¡¡or¡¡subjection¡¡under¡¡which¡¡those¡¡animals¡¡are¡¡to¡¡man¡£¡¡They¡¡do¡¡not¡¡proceed¡¡far¡¡in¡¡half¡­domesticated¡¡species¡£¡¡In¡¡the¡¡cat£»¡¡for¡¡example£»¡¡a¡¡softer¡¡or¡¡harsher¡¡fur£»¡¡more¡¡brilliant¡¡or¡¡more¡¡varied¡¡colors£»¡¡greater¡¡or¡¡less¡¡sizethese¡¡form¡¡the¡¡whole¡¡extent¡¡of¡¡variety¡¡in¡¡the¡¡species£»¡¡the¡¡skeleton¡¡of¡¡the¡¡cat¡¡of¡¡Angora¡¡differs¡¡in¡¡no¡¡regular¡¡and¡¡constant¡¡circumstances¡¡from¡¡the¡¡wild¡­cat¡¡of¡¡Europe¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£¡¡¡£

The¡¡most¡¡remarkable¡¡effects¡¡of¡¡the¡¡influence¡¡of¡¡man¡¡are¡¡produced¡¡upon¡¡that¡¡animal¡¡which¡¡he¡¡has¡¡reduced¡¡most¡¡completely¡¡under¡¡subjection¡£¡¡Dogs¡¡have¡¡been¡¡transported¡¡by¡¡mankind¡¡into¡¡every¡¡part¡¡of¡¡the¡¡world¡¡and¡¡have

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